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論文名稱 Title |
台灣奇馬陸科(倍足綱)的分類研究 Taxonomic Study of Paradoxosomatidae (Diplopoda) in Taiwan |
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系所名稱 Department |
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畢業學年期 Year, semester |
語文別 Language |
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學位類別 Degree |
頁數 Number of pages |
211 |
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研究生 Author |
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指導教授 Advisor |
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召集委員 Convenor |
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口試委員 Advisory Committee |
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口試日期 Date of Exam |
2004-01-31 |
繳交日期 Date of Submission |
2004-03-02 |
關鍵字 Keywords |
倍足綱、奇馬陸科、馬陸、台灣 diplopoda, paradoxosomatidae, millipede, Wang |
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統計 Statistics |
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中文摘要 |
中文摘要 台灣馬陸原記錄58種,分屬於六目,其中22種屬於帶馬陸目 (Polydesmida) 的奇馬陸科 (Paradoxosomatidae)。在台灣採集了379隻奇馬陸標本為主,另向國內其他大學及研究機構商借部分標本共321隻,透過文獻、光學顯微鏡觀察外部型態及電子顯微鏡照相,並比較國外標本,發現當中有3種是同種異名:Helicorthomorpha kosingai (Wang, 1958a) 是 H. orthogonal (Silvestri, 1898) 的同種異名, Kronopolites ralphi Wang, 1957b是K. formosanus (Verhoeff, 1939b) 的同種異名, Oxidus circofera (Verhoeff, 1931) 是Orthomorphella pekuensis (Karsch, 1881) 的同種異名,另外訂正了2個新同種異名:Orthomorpha flavomarginata Gressitt, 1941 是 Orthomorphella pekuensis (Karsch, 1881) 的同種異名,Chamberlinius shengmui Wang, 1957b 是C. piceofasciatus (Gressitt, 1941) 的同種 異名,1種為錯誤鑑定:Cawjeekelia nordenskiöldi (Attems, 1909), 所以在過去發表的台灣奇馬陸當中實際有17種,10屬,另有一個新紀錄種:Chondromorpha xanthotricha (Attems, 1898),兩個新屬有三新種: Aposigipinius gen. n. 有兩新種aviformis sp. n. 及sierwaldae sp. n.,Changius gen. n. 有一新種jhongshan sp. n.,及其他屬10新種:Aponedyopus shenghaii sp. n.,A. sergeii sp. n.,Chamberlinius parhualienensis sp. n.,C. parpiceofasciatus sp. n.,Inversispina taiwanspiralis sp. n. ,Nedyopus latus sp. n.,N. longitudinalis sp. n.,N. microps sp. n.,N. passericaudatus sp. n.,及N. wui sp. n.,所以到目前為止台灣奇馬陸科應有31種,14屬,而本研究中採集到並鑑定出24種13屬奇馬陸。 |
Abstract |
Abstract In previous taxonomic and fannal studies of millipedes for Taiwan, a total of 58 species were recognizied and in 6 orders, in which 22 were in Paradoxosomatidae. We collected about 379 specimens from Taiwan, and borrowed 321 specimens from other universities and research institutions in Taiwan. By referring to literatures, making use of the optics microscope to observe the external patterns of specimens, and taking photos of SEM, and comparing with the holotypes and paratypes loaned from foreign museums, and finding three species are synonyms: Helicorthomorpha kosingai (Wang, 1958a) is a synonym of H. orthogonal (Silvestri, 1898), Kronopolites ralphi Wang, 1957b is a synonym of K. formosanus (Verhoeff, 1939b), and Oxidus circofera (Verhoeff, 1931) is a synonym of Orthomorphella pekuensis (Karsch, 1881), two species are new synonyms: Orthomorpha flavomarginata Gressitt, 1941 is a new synonym of Orthomorphella pekuensis (Karsch, 1881), Chamberlinius shengmui Wang, 1957b is a new synonym of C. piceofasciatus (Gressitt, 1941), one species, Cawjeekelia nordenskiöldi (Attems, 1909), should be misidentified by Wang which should not distribute in Taiwan. Therefore, there are seventeen species in ten genera of Paradoxosomatidae in Taiwan had been recognizied. In addition, a new recorded species: Chondromorpha xanthotricha (Attems, 1898), two new genera contain three new species: Aposigipinius gen. n., contains two new species, aviformis sp. n. and sierwaldae sp. n., Changius gen. n. contains one new species, jhongshan sp. n., besides, anther ten new species belonging to different genera: Aponedyopus shenghaii sp. n., A. sergeii sp. n., Chamberlinius parhualienensis sp. n., C. parpiceofasciatus sp. n., Inversispina taiwanspiralis sp. n., Nedyopus latus sp. n., N. longitudinalis sp. n., N. microps sp. n., N. passericaudatus sp. n., and N. wui sp. n. So, up to now, there are thirty-one species in fourteen genera of Paradoxosomatidae in Taiwan, and in which twenty-four species in thirteen genera are re-collected and verified in this study. |
目次 Table of Contents |
Index Abstract…..…………………………………………………………………….…...1 Introduction………..………………………………………………………………2 Historical review of Family Paradoxosomatidae in Taiwan..........................................2 Millipedes……………………………………………………………………………...3 Family Paradoxosomatidae............................................................................................5 Morphology........................................................................................................5 Systematics…………………………………………………………………….6 Materials and Methods………………….…..…………………………………..8 Where to find the millipedes..........................................................................................8 Collection.......................................................................................................................9 Preservation…………………………………………………………………………..11 SEM photos…………………………………………………………………………..12 Morphological characters.............................................................................................13 Specimens loaned from different Collections..............................................................13 Taxonomic accounts…………………………………………………………….15 Key to families of order Polydesmida in Taiwan…………………………………….15 Key to genera of Paradoxosomatidae in Taiwan..........................................................16 Genus Aponedyopus Verhoeff, 1939似腹馬陸屬……………………………………19 Aponedyopus jeanae (Wang, 1957) 珍氏似腹馬陸 ……………...………….20 Aponedyopus reesi Wang, 1957瑞氏似腹馬陸……………………………….22 Aponedyopus shenghaii sp. n. 聲海似腹馬陸..................................................26 Aponedyopus sergeii sp. n. 石氏似腹馬陸……………………………………29 Key to species of Aponedyopus in Taiwan..........................................................30 Genus Aposigipinius gen. n. 似萎馬陸屬.................................................................. 32 Aposigipinius aviformis sp. n.鳥狀似萎馬陸…………………………….…....33 Aposigipinius sierwaldae sp. n.史氏似萎馬陸………………………….….......36 Genus Cawjeekelia Golovatch, 1980 傑可馬陸屬.....................................................38 Cawjeekelia kanoi (Takakuwa, 1943) 鹿野傑可馬陸........................................39 Cawjeekelia nordenskiöldi (Attems, 1909) 諾氏傑可馬陸............................... 43 Genus Chamberlinius Wang, 1956 章氏馬陸屬…………………………………….46 Chamberlinius hualienensis Wang, 1956花蓮章氏馬陸………………………47 Chamberlinius parhualienensis sp. n. 似花蓮章氏馬陸………………………51 Chamberlinius parpiceofasciatus sp. n. 似深紋章氏馬陸…………………….53 Chamberlinius piceofasciatus (Gressitt, 1941) 深紋章氏馬陸……......………54 Key to species of Chamberlinius in Taiwan.........................................................57 Genus Changius gen. n. 張氏馬陸屬……………………………………………….59 Changius jhongshan sp. n. 中山張氏馬陸…………………………………….60 Genus Chondromorpha Silvestri, 1897 瘤馬陸屬………………………………….63 Chondromorpha xanthotricha (Attems, 1898) 金毛瘤馬陸……….…………..64 Genus Helicorthomorpha Attems, 1914繞馬陸屬......................................................68 Helicorthomorpha holstii (Pocock, 1895) 霍氏繞馬陸......................................70 Genus Inversispina Zhang, 1997 旋馬陸屬................................................................74 Inversispina taiwanspiralis sp. n. 台灣螺旋旋馬陸…………………….…....75 Genus Kronopolites Attems, 1914 攏馬陸屬..............................................................78 Kronopolites formosanus (Verhoeff, 1939b) 台灣攏馬陸..................................79 Genus Nedyopus Attems, 1914 腹馬陸屬……………………………………….….83 Nedyopus latus sp. n. 寬腹馬陸………………………………………….........85 Nedyopus longitudinalis sp. n. 長腹馬陸………………………………….......88 Nedyopus microps sp. n. 小腹馬陸…………………………………………....93 Nedyopus passericaudatus sp. n. 燕尾腹馬陸………………………..….........96 Nedyopus patrioticus (Attems, 1898) 愛國者腹馬陸……………………..... 101 Nedyopus wui sp. n. 吳氏腹馬陸……………………………………….……104 Key to species of Nedyopus in Taiwan...............................................................108 Genus Orthomorpha Bollman, 1893 直形馬陸屬……………………………...….109 Orthomorpha coarctata (Saussure, 1860) 粗直形馬陸....................................110 Genus Orthomorphella Hoffman, 1963 小直形馬陸屬………………………..….116 Orthomorphella pekuensis (Karsch, 1881) 北京小直形馬陸…………….….117 Genus Oxidus Cook, 1911 酸帶馬陸屬……………………………………………122 Oxidus gracilis (Koch, 1847) 雅酸帶馬陸.......................................................123 Discussion...............................................................................................................130 Literature cited.....................................................................................................132 Characteristics of millipedes for identifications (1-3)............................139 Figures list of species Figs. 2.1-2.8, 28.1a-28.2b Aponedyopus jeanae (Wang, 1957).................142, 177 Figs. 3.1-3.8, 29.1a-29.2c Aponedyopus reesi Wang, 1957.......................143, 176 Figs. 4.1-4.9, 30.1a-30.4 Aponedyopus shenghaii sp. n…………….....…144, 177 Figs. 5.1-5.9, 31.1-31.3b Aponedyopus sergeii sp. n................................. 146, 178 Figs. 6.1-6.10, 32.1a-32.4b Aposigipinius aviformis sp. n. .......................148, 179 Figs. 7.1-7.10, 33.1-33.3b Aposigipinius sierwaldae sp. n........................149, 180 Figs. 8.1-8.9, 34.1-34.4 Cawjeekelia kanoi (Takakuwa, 1943)..................151, 181 Figs. 9.1-9.8, 35.1a-35.2 Chamberlinius hualienensis Wang, 1956.........153, 182 Figs. 10.1-10.7, 36.1a-36.1b Chamberlinius parhualienensis sp. n...........154, 182 Figs. 11.1-11.7, 37.1a-37.2 Chamberlinius parpiceofasciatus sp. n..........155, 183 Figs.12.1-12.7, 38.1a-38.2 Chamberlinius piceofasciatus (Gressitt, 1941)............. ............................................................................................................156, 183 Figs.12.8, 39.1-39.2 Chamberlinius shengmui Wang, 1957......................156, 181 Figs. 13.1-13.8, 40.1-40.4 Changius jhongshan sp. n................................157, 184 Figs. 14.1-14.9, 41.1-41.4b Chondromorpha xanthotricha (Attems, 1898)............ ........................................................................................................158, 184 Figs.15.1-15.8, 42.1-42.4 Helicorthomorpha holstii (Pocock, 1895)........159, 185 Figs. 16.1-16.8, 43.1a-43.4 Inversispina taiwanspiralis sp. n...................160, 186 Figs.17.1-17.8, 44.1a-44.4 Kronopolites formosanus (Verhoeff, 1939b)...162, 187 Figs.19.1-19.8, 45.1a-45.3b Nedyopus latus sp. n.....................................164, 188 Figs. 20.1a-20.8, 46.1a-46.3 Nedyopus longitudinalis sp. n......................165, 188 Figs. 21.1-21.8, 47.1a-47.3 Nedyopus microps sp. n..................................167, 189 Figs. 22.1a-22.8, 48.1a-48.4 Nedyopus passericaudatus sp. n…………..168, 190 Figs.18.1-18.6 Nedyopus patrioticus (Attems, 1898)........................................163 Figs. 23.1-23.8, 49.1a-49.4 Nedyopus wui sp. n........................................169, 191 Figs. 24.1-24.7, 50.1-50.3b Orthomorpha coarctata (Saussure, 1860).....171, 191 Figs. 25.1a-25.7, 51.1a-51.1b Orthomorphella pekuensis (Karsch, 1881).............. ....................................................................................................................172, 192 Figs. 26.1a-26.6 Orthomorphella flavomarginata Gressitt, 1941......................173 Figs. 27.1a-27.7, 52.1a-52.2b Oxidus gracilis (Koch, 1847).....................174, 192 Distribution maps of species The siteds of collection of Family Paradoxosomatidae in Taiwan.....................193 The distribution of genus Aponedyopus Verhoeff, 1939 in Taiwan………….194 The distribution of genus Aposigipinius gen. n. in Taiwan…………….…….195 The distribution of Cawjeekelia kanoi (Takakuwa, 1943) in Taiwan................196 The distribution of genus Chamberlinius Wang, 1956 in Taiwan………….....197 The distribution of Changius jhongshan sp. n. in Taiwan…….………………198 The distribution of Chondromorpha xanthotricha (Attems, 1898) in Taiwan...199 The distribution of Helicorthomorpha holstii (Pocock, 1895) in Taiwan..........200 The distribution of Inversispina taiwanspiralis sp. n. in Taiwan.......................201 The distribution of Kronopolites formosanus (Verhoeff, 1939b) in Taiwan…..202 The distribution of genus Nedyopus Attems, 1914 in Taiwan.........................203 The distribution of Orthomorpha coarctata (Saussure, 1860) in Taiwan..........204 The distribution of Orthomorphella pekuensis (Karsch, 1881) in Taiwan.........205 The distribution of Oxidus gracilis (Koch, 1847) in Taiwan.............................206 Index to tables Table 1. A checklist of Paradoxosomatidae from Taiwan………………….…..207 Table 2. Synonyms list ......................................................................................209 Table 3. Species in Wang’s 1964 fauna haven’t been collected……………….21 |
參考文獻 References |
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