蹄蓋蕨屬植物主要分布於亞洲溫帶地區，全球約有220種。本研究根據型態特徵作為蹄蓋蕨屬植物屬下分類架構以及物種分類修訂的依據，台灣蹄蓋蕨屬植物共計有24種2變種，可將其歸屬為五組：耳狀蹄蓋蕨組（Sect. Polystichoides）、硬刺蹄蓋蕨組（Sect. Echinoathyrium）、刺蹄蓋蕨組（Sect. Strigoathyrium）、大葉假冷蕨組（Sect. Atkinsonii）以及日本蹄蓋蕨組（Sect. Niponica）。
根據目前已公開蹄蓋蕨屬植物葉綠體DNA片段（trnL-F以及rbcL），並加入本研究取材自台灣的資料，進行分子親緣關係分析，採用最大簡約法以及貝葉氏導出式分析個別片段：trnL-F區域長度846bp，41種8變種72 OTUs；rbcL區域長度1188bp，共計42種6變種70 OTUs、以及合併序列分析：38種4變種的結果，均支持台灣產蹄蓋蕨類植物所屬的五組，以及台灣不產的蹄蓋蕨組（Sect. Athyrium）以及禾稈蹄蓋蕨組（Sect. Yokoscentia），分別為單系群，支持本研究所提出蹄蓋蕨屬下區分為七組的屬下分類架構。
本文提供台灣產蹄蓋蕨屬植物物種檢索表，以及各物種學名資料、形態描述、引證標本以及分類註記。

The genus Athyrium composed by 220 species in the world, mainly distributed in the Asia temperate regions. According to the observations of the morphological characters, this study revised the taxonomy and recognized 24 species and 2 varieties in Taiwan. The infrageneric scheme also due to morphology, these Taiwanese species were placed into 5 sections: Sect. Polystichoides, Sect. Echinoathyrium, Sect. Strigoathyrium, Sect. Atkinsonii and Sect. Niponica.
Basing on the public DNA sequences data (trnL-F and rbcL), including to this study providing, the molecular phylogeny analysis was state in this study. The Maximum Parsimony and Bayesian inference were used in each fragment: trnL-F region length 846 bp, 72 OTUs in 41 species 8 varieties; rbcL 1188 bp, 70 OTUs in 42 species 6 varieties, and combined data: 42 OTUs in 38 species 4 varieties. The phylograms contributed by each datasets are support the infrageneric scheme provided in this study, 5 sections occuring in Taiwan and 2 other sections (Sect. Athyrium and Sect. Yokoscentia) are monophyletic.
Herein, the key to the species of the Taiwanese Athyrium, science names, descriptions, voucher specimens and notes of each taxa are provided.

壹、前言 1
貳、分類簡史 3
一、屬的確立與分類歷史 3
二、蹄蓋蕨屬植物物種的分類研究 6
三、屬下分類架構 9
四、分子親緣關係 12
五、台灣蹄蓋蕨屬植物的分類研究 13
六、台灣產蹄蓋蕨屬分類架構及物種名錄 15
參、分類特徵評述 17
肆、分子親緣關係研究 22
伍、分類處理 31
Key to the Athyrium Sections and Species in Taiwan 31
耳狀蹄蓋蕨組 Sect. Polystichoides Ching & Y.T. Hsieh 35
1. Athyrium anisopterum Christ 宿蹄蓋蕨 36
2. Athyrium nakanoi Makino 紅苞蹄蓋蕨 39
3. Athyrium puncticaule (Bl.) T. Moore 密腺蹄蓋蕨 41
4. Athyrium takeoi (Hayata) Tagawa 細裂蹄蓋蕨 44
硬刺蹄蓋蕨組 Sect. Echinoathyrium Ching & Y.T. Hsieh 48
5. Athyrium minimum Ching 小蹄蓋蕨（七星山蹄蓋蕨） 49
6. Athyrium oppositipennum Hayata 對生蹄蓋蕨 51
6.1 Athyrium oppositipennum var. oppositipennum對生蹄蓋蕨 51
6.2 Athyrium oppositipennum var. pubecent (Tagawa) Liu 逆羽蹄蓋蕨 53
7. Athyrium tripinnatum Tagawa 三回蹄蓋蕨 54
8. Athyrium vidalii (Franch. & Sav.) Nakai 山蹄蓋蕨 56
刺蹄蓋蕨組 Sect. Strigoathyrium Ching et Y.T. Hsieh 59
9. Athyrium arisanense (Hayata) Tagawa阿里山蹄蓋蕨 61
10. Athyrium auriculatum Seriz. 耳垂蹄蓋蕨 64
11. Athyrium cryptogrammoides Hayata 合歡山蹄蓋蕨 66
12. Athyrium delavayii H. Christ 溪谷蹄蓋蕨 68
12.1 Athyrium delavayi var. dalavayi 溪谷蹄蓋蕨 68
12.2 Athyrium delavayi var. subrigescens (Hayata) Liu 姬蹄蓋蕨 69
13. Athyrium epirachis (Christ) Ching 軸果蹄蓋蕨 71
14. Athyrium erythropodum Hayata 紅柄蹄蓋蕨 72
15. Athyrium iseanum Rosenst. 細葉蹄蓋蕨 75
16. Athyrium leiopodum (Hayata) Tagawa 小葉蹄蓋蕨 77
17. Athyrium nigripes (Blume) T. Moore 蓬萊蹄蓋蕨 78
18. Athyrium otophorum (Miq.) Koidz. 光蹄蓋蕨 80
19. Athyrium pubicostatum Ching et Z. Y. Liu 假軸果蹄蓋蕨 81
20. Athyrium silvicola Tagawa 高山蹄蓋蕨 83
21. Athyrium strigillosum (T. Moore ex E.J. Lowe) Salomon 生芽蹄蓋蕨 85
22. Athyrium yui Ching 俞氏蹄蓋蕨 86
大葉假冷蕨組 Sect. Atkinsonii Liu 89
23. Athyrium atkinsonii Bedd. 大葉假冷蕨 89
日本蹄蓋蕨組 Sect. Niponica Ching et Y.T. Hsieh 92
24. Athyrium niponicum (Mett.) Hance 日本蹄蓋蕨 92
陸、引用文獻 94

表1台灣產蹄蓋蕨屬植物學名變革 100
表4.1 分子譜系分析材料對照表 101
表4.2 分子序列實驗用引子 104
表4.3 譜系分析統計表 105
圖2.1 Athyrioides phylogeny basing on rbcL, Sano et al, 2002 106
圖2.2 Athyriaceae sensu Ching, basing on trnL-F region, Wang, et al., 2003 107
圖3.1. 蹄蓋蕨類根莖型 108
圖3.2蹄蓋蕨類近軸面刺的類型及相關位置 108
圖4.1 MP Phylogram of Athyrium basing trnL-F region 109
圖4.2 Athyrium phylogram basing on trnL-F Bayesian inference analysis 110
圖4.3 MP Phylogram of Athyrium basing rbcL region 111
圖4.4 Athyrium phylogram basing on rbcL Bayesian inference analysis 112
圖4.5 MP Phylogram of Athyrium basing on trnL-F and rbcL region 113
圖4.6 Athyrium phylogram basing on trnL-F and rbcL Bayesian inference analysis 114
附錄一　Name List of Genus Athyrium in the World 115

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