植物系統分類本在追求內含親緣關係架構之分類系統，為達此一目標，我們必須藉助各類可資利用的特徵來建構它。蕨類植物之屬間劃分一直存在許多問題，許多的特徵亦值得重新去檢視，如毛被物，早被蕨類植物學家認為重要之分類特徵一；但臺灣對此主題之研究甚少，故本研究欲以毛被物微細特徵，嘗試釐清臺灣產廣義鱗毛蕨科之屬間關係。觀察重點在於葉面上分佈之毛與葉柄基部之鱗片，結果，可將葉面毛的部份分為：單細胞毛、棒狀單細胞腺體、單列細胞毛、肉刺毛、倒伏狀腺毛、單列細胞腺毛、疣狀物；而葉柄基部鱗片部份，可依其邊緣、顏色分佈、分支向度等特性分為：全緣、鋸齒緣、單細胞分支、單列細胞分支、多細胞腺體分支、背腹性生長分支等。藉由毛被物的研究，我們在肉刺蕨屬、柄囊蕨屬、耳蕨屬、擬鱗毛蕨屬、肋毛蕨屬、三叉蕨屬、蹄蓋蕨屬、冷蕨屬、亮毛蕨、岩蕨屬得到良好的分群結果；但另一方面，雙蓋蕨屬和鱗毛蕨屬，結果顯示其毛被物特徵異質性高，且無法與在今之系統相互對應，有待進一步廣泛之研究。

A unifying goal of plant systematics is in pursuit of a natural system that means phylpgenetic relationships. We can construct it by means of various characters. There are many arguments about generic circumscription of Pteridophyta. It’s worth while to reexamine some characters, such as trichome. Trichomes have long been considered one of the most important characters by pteridologists. However, The study of trichomes in Taiwan is very rare. In the present study, trichome morphology is used to assess phylogenetic relationships among genera of Dryopteridaceae (sensu Kramer et al.) . The observation of trichomes focus on hairs on the lamina and scales on the base of stipes. Lamina surface hairs are classified into unicellular hairs, club-like unicellular glands, uniseriate hairs, spine-like hairs, appressed glandular hairs, uniseriate hairs with a glandular head and verruca. Stipe base scale margins are classified into entire, serrate, unicellular branch, uniseriate branch, multicellular branch with a glandular head, and dorsi-ventral branch. Color distribution and branch dimension of scales were also good differentiation characters. The main taxonomic conclusions are as follows: (1)Trichome characters support the distinctness of Nothoperanema, Peranema, Polystichm, Dryopsis, Ctenitis, Tectaria, Athyrium, Cystopteris, Acystopteris, and Woodsia; (2) Diplazium and Dryopteris are hererogenous and show little correaltion to exist system. The latter two genera need further research.

目次


摘要 I

Abstract II

目次 III

表目次 IV

圖目次 V

前言 1

材料與方法 3

結果與討論 4

毛被物特徵 4

葉面毛之部份 6

葉柄基部鱗片之部分 10

屬上階層之系統分類應用 16

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