本論文探討台灣產鴨嘴草屬 ( Ischaemum ) 植物之分類，主要依據植株習性、莖桿、葉、花序、小穗、花及穎果等形態特徵。研究的結果將台灣產鴨嘴草屬植物區分為5種，即芒穗鴨嘴草 ( Ischaemum aristatum L. var. aristatum ) (另將毛穗鴨嘴草I. aristatum var. momiyamai (Honda) Hsu 併入本種)、瘤鴨嘴草 ( I. barbatum Retzius ) (將瘤鴨嘴草之一變種 I. barbatum var. gibbum (Trin.) Ohwi 併入)、細毛鴨嘴草 ( I. ciliare Retzius ) (將印度鴨嘴草 I. indicum (Houtt.) Merr. 處理為本種之同物異名)、田間鴨嘴草 ( I. rugosum Salisbury ) (將屏東鴨嘴草 I. akonense Honda 及田間鴨嘴草之一變種 I. rugosum var. segetum Hackel 均併入) 及小黃金鴨嘴草 ( I. setaceum Honda )。因未見標本，本研究將鴨嘴草 ( I. aristatum var. crassipes (Steud.) Yonekura )、黃金鴨嘴草 ( I. aureum (Hook. & Arn.) Hackel )、無芒鴨嘴草 ( I. muticum L. ) 和帝汶鴨嘴草 ( I. timorense Kunth ) 皆暫列為疑問種。另外，研究結果顯示小穗之芒、壓扁情形(背腹壓扁或兩側壓扁)及下位外穎等，是區別台灣產本屬植物的重要特徵。本研究詳細描述各種之形態特徵並作分種檢索表。此外，並記載各種之分布及產地。

A taxonomic study on Ischaemum of Taiwan has been carried out through the past three years, according to morphological characters including habit, culm, leaf, inflorescence, spikelet, flower and caryopsis. The results indicated that there are totally 5 species of the genus occurring in Taiwan, i. e. Ischaemum aristatum L. var. aristatum (including I. aristatum var. momiyamai (Honda) Hsu) , I. barbatum Retzius (including I. barbatum var. gibbum (Trin.) Ohwi), I. ciliare Retzius (including I. indicum (Houtt.) Merr.), I. rugosum Salisbury (including I. akonense Honda and I. rugosum var. segetum Hackel), and I. setaceum Honda. Because of no specimens available for examination in the herbariums of Taiwan, I. aristatum var. crassipes (Steud.) Yonekura, I. aureum (Hook. & Arn.), I. muticum L., and I. timorense Kunth are treated as doubtful species. In addition, the features of awn, compression and lower glume of spikelets are realized the diagnostic characters to distinguish the species. In this study, the detailed descriptions of and a key to the species and varieties are given. Moreover, the distribution and habitat of each species are also provided.

壹、前言 ...................................................................... 1
貳、文獻探討 .............................................................. 4
參、材料與方法......................................................... 23
肆、結果與討論
一、台灣產鴨嘴草屬植物之形態特徵 .................. 24
二、台灣產鴨嘴草屬植物之地理分布 .................. 48
三、台灣產鴨嘴草屬植物之分類處理 .................. 57
伍、結論 .................................................................. 116
陸、參考文獻 .......................................................... 117
柒、附錄 .................................................................. 123

Backer, A. C.& Bakhuizen R. C. 1968. Fl. Java 3, pp. 591-593. Wolters-Noordhoff N.V., Groningen.
Bor, N. L. 1960. The Grasses of Burma, Ceylon, India and Pakistan (excluding Bambuseae), pp. 180-182. Pergamon Press, London and N.Y.
Chapman, G. P. 1996. The Biology of Grass, pp. 14-35. CAB International, Wallingford.
Chen, C. C. & Hsu, C. C. 1962. Cytological studies on Taiwan grasses (2). Chromosome numbers of some misecellaneous tribes, J. Jap. Bot. 37(10): 300-313.
Chen, S. L. 1997. Flora of China 10(2), pp. 154-165. Science Press, Peking.
Cheng, W. T. 2000. The Illustrated Flora of Taiwan, pp. 1759-1763. 茂昌圖書, Taipei.
Clayton, W. D., Harman, K. T. & Williamson, H. GrassBase - The Online World Grass Flora. Retrieved November 18, 2006 from the World Wide Web:
http://www.kew.org/data/grasses-db/ident.htm
FAO Cropard Grassland Service (AGPC). Retrieved January 18, 2007 from the World Wide Web:
http://www.fao.org/AG/AGP/AGPC/doc/Default.htm
Flora of Australia Online (1993). Retrieved February 7, 2007 from the World Wide Web:
http://www.anbg.gov.au/abrs/online-resources/flora
Fujimoto, Y. 1967. Classification of Ischaemum Linn. based on vegetative characters, especially on the ligule, Hikobia. 5. 104-111.
Gamble, J. S. & Fischer, C.E.C. 1934. Flora of Madras, pp. 1717-1723.
Hackel, E. 1889. Monographiae Phanerogamarum 6: 200-253.
____. 1899. Enumeratio Graminum Japoniae, Bulletin de L’Herbier Boissier. 7(10): 723.
____. 1904. Supplementa Enumerationis Graminum, Japonle, Formosae, Coreae, Bulletin de L’Herbier Boissier ser.II 4: 527.
Hatusima, S. 1971. Flora of the Ryukyus, pp. 709-711.
Hayata, B. 1918. Icones Plantarum Formosanarum 7, pp. 77-78. Bureau of Productive Industries, Taipei.
Henry, A. 1896. A List of Plants from Formosa, Trans. Asiatic Soc. Jap. 24 (Supplement), pp. 108. R. Meiklejohn & Co., Yokohama.
Hideaki, O., Shinobu, A. & Gerard, T. 2005. Miquel’s new taxa of the vascular plants described from Japan in Prolusio Florae Japonicae and some other works, Museum Univ. Tokyo Bulletin 41. Retrieved March 11, 2007 from the World Wide Web:
http://www.um.u-tokyo.ac.jp
Hooker, W. J. & Arnott, G. A. 1841. Botany of Captain Beechey's Voyage, pp. 273-274.
Honda, M. 1923. 日本產かものはし屬並わたがゐ屬就テ, Bot. Mag. Tokyo 37: 120-124.
____. 1924. Revisio Graminum Japoniae IV, Bot. Mag. Tokyo 38: 49-56.
____. 1927. Revisio Graminum Japoniae XII, Bot. Mag. Tokyo 41: 8.
____. 1927. Revisio Graminum Japoniae XIII, Bot. Mag. Tokyo 41: 377-379.
____. 1930. Monographia Poacearum Japonicarum, Bambusoideis exclusis, J. Fac. Sci. Univ. Tokyo III 3: 350-366.
Hsieh, A. T. & Yang, T. I. 1969. Nomenclature of Plants in Taiwan, pp. 145-147. College of Agriculture NTU, Taipei.
Hsu, C. C. 1962. Notes on the Grasses of Formosa, J. Jap. Bot. 37(9): 275-279.
____. 1963. A study on Formosan Panicum(Gramineae), with special reference to their lodicules, J. Jap. Bot. 38(3): 75-86.
____. 1971. A Guide to the Taiwan Grasses, with keys to subfamilies, tribes, genera and species, Taiwania 16: 322-324.
____. 1975. Taiwan Grssses, pp. 1-884. Taiwan Prov. Educ. Assoc., Taipei.
____. 1978. Flora of Taiwan 5: 663-671.
____. 2003. Flora of Taiwan 2nd ed. 6: 554-562.
Huang, T. C. 2002. 植物命名指南, pp. The Council of Agriculture, Taipei.
Huang, T. C. & Hsiao, C. L. 2003. Checklist of Taiwan Vascular Plants, pp. 142. 南天書局, Taipei.
Kawakami, T. 1910. A List of Plants of Formosa, pp. 140. Bureau of Productive Industries, Taipei.
Koyama, T. 1962. Nomenclatorial remarks on some grasses, J. Jap. Bot. 37(8): 238-239.
____. 1987. Grass of Japan and its Neighboring Region, pp. 454-465. Kodansha, Tokyo.
Kunth, C. S. 1830. Revision des Graminees, pp. 369-372.
Linnaeus, C. 1753. Species Plantarum 2: 1049.
Liu, H. Y., Yang, Y. P., Kuo, C. S. & Chen, C. H. 2001. Maunal of Taiwan Vascular Plants 5, pp. 114-146. The Council of Agriculture, Taipei.
Makino, T. & Nemoto, K. 1925. Flora of Japan, pp. 1457-1460. Nipponshokubutsusoran Publishing Association, Tokyo.
____. 1931. Flora of Japan 2nd ed., pp. 1352-1355. Shunyodo Shoten Nihonbashi Tori Sanchome, Tokyo.
Mary E. B. Grass Treatments for North American. Retrieved February 6, 2005 from the World Wide Web:
http://herbarium.usu.edu/treatments/Ischaemum.htm
Masamune, G. 1936. Short Flora of Formosa, pp. 236-237. Kudoa, Taipei.
____. 1954. A List of Vascular Plants of Taiwan, pp. 146-147.
Matsumura, J. 1905. Index Plantarum Japonicarum 2: 60-62.
Matsumura, J. & Hayata, B. 1906. Enumeratio Plantarum Formosanarum, Journal College of Science Imp. Univ. Tokyo 22: 525-527.
Merrill, E. D. 1925. An Enumeration of Philippine Flowering Plants, pp. 37-39., Bureau of Printing, Manila.
Merrill, E. D. 1938. A critical consideration of Houttuyn’s new genera and new species of plants, 1773-1783, Journal of the Arnold Arboretum 19(4): 320. 1938.
Nakai, T. 1919. Notula ad plantas Japonia et Corea XIX, Bot. Mag. Tokyo 33: 1-3.
Natural History Museum, London. 2007. Linnaean Plant Name Typification Project. Retrieved June 11, 2007 from the World Wide Web:http://internet.nhm.ac.uk/jdsml/research-curation/projects/linnaean-typification
Nemoto, K. 1936. Flora of Japan Supplement, pp. 934-936. Shunyodo Shoten Nihonbashi Tori Sanchome, Tokyo.
Ohwi, J. 1942. Gramina Japonica IV, Acta Phytotaxonomica et Geobotanica 11(3): 171-175.
____. 1965. Flora of Japan (in English), pp. 193-194. Smithsonian Institution Press, Washington, D.C.
Pacific Island Ecosystems at Risk (PIER). Retrieved January 19, 2007 from the World Wide Web:
http://www.hear.org/pier
Ravi, N.,N. Mohanan, T. Shaju, M. S. Kiran Raj, and R. Rajesh, 2001. Three more new species of Ischaemum L. (Poaceae) from Kerala, India. Bot. Bull. Acad. Sin. 42: 223-230.
Rendle, A. B. 1904. An enumeration of all the plants known from China Proper, Formosa, Hainan, Corea, the Luchu Archipelago, and the Island of Hongkong, together with their distribution and synonymy, Linn. Soc. J. Bot. 36: 364-366.
Retzius, A. J. 1791. Observationes Botanicae 6: 35-36.
Sasaki, S. 1928. List of Plants of Formosa, pp. 68-69. The Natural History Society of Formosa, Taipei.
Shimabuku, K. 1997. Check List Vascular Flora of the Ryukyu Islands, pp. 668-670. 九州大學出版會.
Spangler, R., Zaitchik, B. & Kellogg, E. 1999. Andropogoneae evolution and generic limits in Sorghum (Poaceae) using ndhF sequences, System Botany 24(2): 267-281.
Steudel, E. G.. 1854. Synopsis Plantarum Glumacearum 1: 375.
Sun, B. S. & Philips, S. M. 2006. Flora of China 22: 609-613.
Thellung A. 1912. Combinationes novae, Feddes Repertorium Specierum Novarum Regni Vegetabilis 10: 289.
Thomas, R. S., Khidir, W. H., Christopher, S. C. & Mary E. B. 1986. Andropogoneae, Grass Sysematic and Evolution, pp. 307-309. Smithsonian Institution Press, Washington, D.C. and London.
Veldkamp, J. F. 1991. Miscellaneous notes on Southeast Asian Gramineae. VI, Blumea 36: 179-181.
W3TRPPICOS. Retrieved November 22, 2006 from the World Wide Web:
http://mobot.mobot.org/W3T/Search/vast.html
Walker, E. H. 1976. Flora of Okinawa and the Southern Ryukyu Islands, pp. 235-236. Smithsonian Institution Press, Washington, D.C.
Watson, L. & Dallwitz. M. J. Grass Genera of the World. Retrieved June 6, 2007 from the World Wide Web:
http://www.biologie.nui-hamburg.de/b-online/delta/grass/www/ischaemum.htm
____. Classification (1998), Grass Genera of the World. Retrieved July 7, 2007 from the World Wide Web:
http://www.biologie.nui-hamburg.de/b-online/delta/grass/www/class.htm
Yao, J. N. 1994. Species List of Vascular Plants of Taiwan, pp. 100. 台大實驗林管理處, 南投竹山.
Yang, T. I. 1973. 台灣植物名彙, pp. 147-148. Published by the auther, Taipei.
____. 1982. A List of Plants in Taiwan, pp. 212-214. 天然書社, Taipei.
Yang, Y. P. 2006. 墾丁國家公園水生植物圖鑑, pp. 141-143. Kending National Park Administration Office, Hengchun.
Yang, Y. P., Yen, S. H. & Lin, C. K. 2001. Illustrated Guide to Aquatic Plants of Taiwan, pp. 142. The Council of Agriculture, Taipei.
Yonekura, K. 2005. Taxonomic notes on vascular plants in Japan and its adjacent regions, J. Jpn. Bot. 80: 332.
Zhao Huiru, New grasses in Chinese Ischaemum L., 1983 Acta Bot. Yunnanica 5(4): 343-354.