旋梯螺在芋螺超科的分類地位不明，曾被歸在旋梯螺科、捲管螺科及芋螺科內，但芋螺超科大而多元，全世界的物種數可能超過一萬種，傳統上以殼形及齒舌形態為分類依據，分為芋螺科、筍螺科、捲管螺科，之後，以前腸的解剖構造、殼形及齒舌形態將三科修訂至七科。本研究以粒線體的16S rDNA片段探討芋螺超科的分類及旋梯螺在超科內的分類位階，結果顯示芋螺超科分為三大系群，推測此三系群分別為芋螺科、筍螺科及捲管螺科；各系群內的平均遺傳距離分別為0.12、0.10及0.10，系群間則在0.14~0.17間；筍螺科及捲管螺科可能為姊妹群或二者在同一類群下，且筍螺科與捲管螺科的親緣關係比芋螺科近。此外，旋梯螺和Bathytoma luhdorfi 雖具有捲管螺的螺旋殼形，其親緣關係卻與芋螺科系群的Conus屬較接近，其他原歸於捲管螺科的三個物種（如Oenopota sagamiana、Phymorhynchus buccinoides 和Raphitoma linearis）亦歸於芋螺科系群。此分類結果不支持Powell (1966) 和 Kohn (1998) 以殼形特徵為依據的傳統芋螺超科分類，而與Taylor等人 (1993)、Rosenberg (1998) 及Bouchet和Rocroi (2005) 以前腸解剖特徵所做的支序親緣結果類似，而芋螺超科中空魚叉狀齒舌及毒腺構造可能由各科獨立演化而產生。

The taxonomic status of the Japanese Wonder Shell, Thatcheria mirabilis is questionable, because it has been classified in the family of Thatcheriidae, Turridae or Conidae (Superfamily: Conoidea). Conoidea is a large and diverse superfamily with more than 10,000 species. Based on shell and radula characters, it is classified into three families, i.e. Conidae, Terebridae and Turridae. However, seven families have been proposed based on foregut structure, shell and radula morphology. In the present study, the molecular phylogeny of Conoidea and the taxonomic status of Thatcheria mirabilis were determined by mitochondria DNA 16S rDNA. The results show that Conoidea includes three clades, presuming Conidae, Terebridae and Turridae. The mean genetic distances within clades were 0.12, 0.10 and 0.10, respectively. And, the distances between clades were 0.14~0.17. Phylogenetic trees reveal that Terebridae and Turridae were within the same group or sister group, Terebridae was closer to Turridae than to Conidae. Although Thatcheria mirabilis and Bathytoma luhdorfi have turrid-form shells, their phylogenetic relationship was close to Conus which was in Conidae`s clade. Some other species, i.e. Oenopota sagamiana、Phymorhynchus buccinoides and Raphitoma linearis were also in Conidae`s clade which had been placed in Turridae. In general, the results are consistent with the cladistic classification by Taylor et al (1993), Rosenberg (1998) and Bouchet & Rocroi (2005), but differenrt from the classification by Powell (1966) and Kohn (1998) based on shell characters. Additionally, the hollow, harpoon-like teeth and venom apparatus in Conoidea might independently evolve in each family.

謝辭.............................................................I
中文摘要....................................................II
英文摘要...................................................III
目錄...........................................................IV
表目錄........................................................V
圖目錄.......................................................VI
附錄目錄..................................................VII
前言............................................................1
材料與方法................................................7
結果..........................................................11
討論..........................................................16
參考文獻..................................................21
表..............................................................26
圖..............................................................35
附錄..........................................................52

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